LSU football defensive back Jardin Gilbert had a costly targeting penalty in Sunday's season-opener against USC.

Gilbert led with the crown of his helmet, hitting Trojans wide receiver Kyron Hudson directly on his helmet after Hudson made a 20-yard catch off a pass from Miller Moss. Hudson made the catch at the 27-yard line with 18 seconds left in the game and the score tied 20-20.

The penalty, however, put the ball at the 13-yard line, with Woody Marks scoring the go-ahead touchdown on the next play. Gilbert was ejected from the game and will face a suspension for the first half of LSU's Week 2 game vs. Nicholls.

REQUIRED READING: USC upsets LSU in statement win

USC's Mason Cobb picked off LSU quarterback Garrett Nussmeier from scrimmage on the next LSU play to pick up the victory for the Trojans.

Here's a look at the costly targeting call against Gilbert, which helped USC defeat LSU on Sunday in Las Vegas:

Look: LSU DB Jardin Gilbert ejected for targeting on USC's Kyron Hudson

With that, LSU starts 0-1 for the third-straight season under Brian Kelly.

This article originally appeared on USA TODAY: Jardin Gilbert targeting call leads to USC game-winning touchdown vs LSU