Michigan football tight end Colston Loveland caught a third-and-8 pass against Oregon to start the fourth quarter right in front of head coach Sherrone Moore, but the play was ruled incomplete.

Fans at Michigan Stadium in Ann Arbor booed, but Moore then challenged the call on the field. Though Loveland got a foot down in bounds, he did not survive the ground, with the ball jarring loose as he hit the sideline around the Michigan 36-yard line in the Wolverines' eventual 38-17 loss.

After a quick review, the call on the field was confirmed, sparking longtime CBS color commentator Gary Danielson to remark:

"That is just a mistake on the Michigan staff. They do not know the rule, and it cost them a timeout.

"They just don't understand the rule."

Michigan lost a key timeout, and punted with 14:49 remaining down 31-17, left with one timeout after having too many men on the field earlier.

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Danielson also said someone in the Michigan press box "upstairs" should have told Moore not to challenge after seeing the replay.

The Wolverines defense stepped up to force a punt, giving Michigan a fresh set of downs at 12:23 to play. The Wolverines drove to the Ducks' 10-yard line, but were stopped on downs after a failed trick play on a throwback to backup quarterback Alex Orji with 7:24 left.

"I just thought Davis Warren earned the throw," Danielson said. "He didn't get it on third down (Kalel Mullings 2-yard run). And I thought the way he played, he earned either the throw on third or the throw on fourth down. That's my opinion."

"I agree with you," play-by-play partner Brad Nessler said.

In the third quarter, Michigan committed a penalty lining up over the long snapper on an odd punt formation, giving the Ducks an automatic first down. Then later in the drive on a second down, substituted on defense and was caught with too many men on the field.

"That one was really on the coaching staff, that was a bad call to make a substitution there," Danielson said.

Oregon earned a field goal on the drive to go up 31-17.

This article originally appeared on Detroit Free Press: Michigan's Sherrone Moore easily loses challenge vs Oregon